Jonathan came to Cambridge in 2005 to read Natural Sciences at Clare College, graduating in 2008. He completed an MPhil in History and Philosophy of Science in 2009 and is currently in the final stages of writing up his PhD dissertation. The dissertation, which lies at the intersection of philosophy and evolutionary biology, examines the conceptual foundations of our current theories of the evolution of cooperation and considers the extent to which the multicellular organism may be regarded as a product of social evolution among free-living cells.

Selected publications:

‘Hamilton’s rule and its discontents’, British Journal for the Philosophy of Science (forthcoming, 2013)

‘On the “simulation argument” and selective scepticism’, Erkenntnis (forthcoming, 2013)

‘Collective action in the fraternal transitions’, Biology and Philosophy 27:363-380 (2012)